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Open front and rear. No Toyota has locking front (I guess you could mod if you are so inclined). FC has a locked rear on his LR4, e.g,, but I doubt he has needed it much. The good thing with Toyota is that there is some aftermarket close to OEM and it seems possible to add a lockable rear through ARB or some such thing.
To get locking rear and stay stock, in the current line-up, you have to give up the center diff and go to 2wd/locked-4wd optionality. Maybe driving a locked 4wd on the highway (4hi or H4) is not so bad in mixed conditions, I'm not sure. Say changing lanes across slush accumulations at 50mph, something a rwd car with snow tires may have some issues with but a subaru or audi powers through. Or slick turns. I worry a locked center will be odd to drive (compared to what I'm used to).
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